MANUSCRIPTS by members of the Math Skills Development Project (MSDP) Team:

This page is being kept updated to include most recent data. Please check back.

*Note: all manuscripts are peer reviewed unless otherwise noted:

# denotes review articles
^ denotes book chapters

Manuscripts on Math or the MSDP:

D Berch & MMM Mazzocco (Eds.), Why is Math So Hard for Some Children: The Nature and Origins of Mathematics Learning Difficulties and Disabilities. Baltimore, MD: Brookes Publishers. (2007).  Listed on Brookes Publishing

MMM Mazzocco & J Ross & (Eds.), Neurogenetic Developmental Disorders: Variation of Manifestation in Childhood. Cambridge, MA: MIT Press. (2007). Listed on MIT Press.

Published Journal Articles

Mazzocco, M. M. M., & Hanich, L. B. (2010). Math achievement, numerical processing, and executive functions in girls with Turner syndrome: Do girls with Turner syndrome have math learning disability? Learning and Individual Differences. (in press).

Murphy, M. M., & Mazzocco, M. M. M. (2009). The trajectory of mathematics skills and working memory thresholds in girls with fragile X syndrome. Cognitive Development, 24, 430-449.

Mazzocco, M. M. M. (2009). Mathematical learning disability in girls with Turner syndrome: A challenge to defining MLD and its subtypes. Developmental Disabilities Research Reviews, 15, 35-44.

Mazzocco, M. M. M. (2009). An Introduction to the Special Issue: Pathways to Mathematical Learning Difficulties and Disabilities. Developmental Disabilities Research Reviews, 15, 1-3.

Dennis, M., Berch, D. B., & Mazzocco, M. M. M. (2009). Mathematical learning disabilities in special populations: Phenotypic variation and cross-disorder comparisons. Developmental Disabilities Research Reviews, 15, 80-89.

Mazzocco, M.M.M. (2008). Introduction: Mathematics Ability, Performance, and Achievement. Developmental Neuropsychology, 33:3, 197-204.

Mazzocco, M.M.M., & Devlin, K.T. (2008). Parts and holes: Gaps in rational number sense in children with vs. without mathematical learning disability. Developmental Science. 11:5, 681-691

Halberda, J., Mazzocco, M. M. M., & Feigenson, L. (2008). Individual differences in nonverbal number acuity correlate with maths achievement. Nature. Published online 7 September, 2008 DOI 10:1038/Nature 07246.

Murphy MM, & Mazzocco MMM. Rote numeric skills may mask underlying mathematical disabilities in girls with fragile X syndrome. Developmental Neurospychology (2008), 33: 345-364.

Mazzocco, M.M.M., Devlin, K.T., & McKenney, J.L. Is it a fact? Timed arithmetic performance of children with mathematical learning disabilities (MLD) varies as a function of how MLD is defined. Developmental Neuropsychology (2008), 33:318-344.

Murphy MM, Mazzocco MM M, Hanich LB, & Early M. Cognitive characteristics of children with mathematics learning disability (MLD) vary as a function of the cut-off criterion used to define MLD. Journal of Learning Disabilities (2007), 458-478.

D Berch & MMM Mazzocco (Eds.), Why is Math Hard for Some Children: The Nature and Origins of Mathematics Learning Difficulties and Disabilities. Baltimore, MD: Brookes Publishers. (2007).

Mazzocco MMM. When a genetic disorder is associated with learning disabilities. In: MMM Mazzocco & J Ross & (Eds.), Neurogenetic Developmental Disorders: Variation of Manifestation in Childhood. MIT Press. (2007). pp 415-436.

Gersten R., Clark B., & Mazzocco MMM. Historical and Contemporary Perspectives on Mathematics Disabilities. In D Berch & MMM Mazzocco (Eds.), Why is Math So Hard for Some Children: The Nature and Origins of Mathematics Learning Difficulties and Disabilities. Brookes Publishers (2007). pp 7-27.

Mazzocco MMM. Issues in Defining Mathematical Learning Disabilities and Difficulties. In D Berch & MMM Mazzocco (Eds.), Why is Math So Hard for Some Children: The Nature and Origins of Mathematics Learning Difficulties and Disabilities. Brookes Publishers (2007). pp 29-47.

Mazzocco MMM. Early predictors of children’s mathematical learning difficulties: Variations in children’s difficulties with math. Exchange (2007); March/April, 40-45.

Mazzocco MMM, & Kover, ST. A longitudinal assessment of the development of executive function and math skills and their association with math performance. Child Neuropsychology (2007): 13:18-45.

Tsui JM, & Mazzocco MMM. Effects of math anxiety and perfectionism on timed versus untimed math testing in mathematically gifted sixth graders. Roeper Review (2007), 29:132-139.

Lachance J, & Mazzocco MMM. A longitudinal analysis of sex differences in math and special skills in primary school age children. Learning & Individual Differences (2006); 16: 195-216.

Mazzocco MMM, Bhatia NS, & Lesniak-Karpiak K. Visuospatial skills and their association with math performance in girls with fragile X or Turner Syndrome. Child Neuropsychology (2006); 12: 87-110.

Murphy MM, Mazzocco MMM, Gerner G, & Henry AE. Mathematics learning disability in girls with Turner Syndrome or fragile X Syndrome. Brain and Cognition (2006); 61: 195-210.

Garrett AJ, Mazzocco MMM, Baker L. Development of the metacognitive skills of prediction and evaluation in children with or without math disability. Learning Disabilities Research and Practice (2006); 21: 77-88.

Russell H F, Wallis D, Mazzocco MMM, Moshang T, Zackai E, Zinn A, Ross JE, & Muenke M. Increased prevalence of Attention-Deficit/Hyperactivity Disorder (ADHD) in girls with Turner syndrome with no evidence of an imprinting effect for cognitive performance or ADHD. Pediatric Psychology (2006); 31: 945-955.

Mazzocco MMM, Thompson L, Sudhalter V, Belser RC, Lesniak-Karpiak K, & Ross JL. Language use in females with fragile X or Turner syndrome during initial social interactions. Journal of Developmental and Behavioral Pediatrics (2006) 27: 319-328.

Kirk J, Mazzocco MMM, & Kover, ST. Assessing executive dysfunction in girls with fragile X or Turner syndrome using the Contingency Naming Test. Developmental Neuropsychology. (2005); 28(3): 755-777.

^ Mazzocco MMM, & McCloskey M. Math performance in girls with Turner or Fragile X syndrome. In Campbell, J (ed), Handbook of Mathematical Cognition. (2005); Hove, East Sussex, Great Britain: Psychology Press. pp 269 -297.

Mazzocco MMM. Challenges in identifying target skills for math disability screening and intervention. Journal of Learning Disabilities. (2005); 38: 318-323.

Mazzocco MMM, & Thompson RE. Kindergarten predictors of math learning disability. Learning Disabilities Research and Practice. (2005); 20: 142-155.

Mazzocco MMM, & Gerner GG. Genetic disorders and learning disability. Perspectives. (2004); 30(3): 18-23.

Mazzocco MMM & Myers GF. Complexities in identifying and defining mathematics learning disability in the primary school age years. Annals of Dyslexia. (2003); 53: 218-253.

Mazzocco MMM, Myers GF, Thompson LA, & Desai SS. Possible explanations for children's literal interpretations of homonyms. Journal of Child Language. (2003); 30(4): 879-904.

Mazzocco MMM, & Myers GF. Maximizing enrollment efficiency for school-based education research. Journal of Applied Social Psychology. (2002); 32:1577-1587.

Mazzocco MMM. Math learning disability and math LD subtypes: Evidence from studies of Turner syndrome, fragile X syndrome, and neurofibromatosis type 1. Journal of Learning Disabilities. (2001); 34(6): 520-533.

Mazzocco MMM & Kelley R. Preliminary evidence for a cognitive phenotype in Barth syndrome. American Journal of Medical Genetics. (2001); 102: 372-378.

Teisl JT, Mazzocco MMM, & Myers GF. Assessing the utility of kindergarten teacher ratings for predicting first grade academic achievement. Journal of Learning Disabilities. (2001); 34(3) 286-293.

Mazzocco MMM. Developmental changes in indicators that literal interpretation of homonyms are associated with conflict. Journal of Child Language. (1999); 26: 393-417.

Mazzocco MMM, Myers GF, Harum KH, & Reiss AL. Children's participation in genetic prevalence research: Influences on enrollment and reports of parent satisfaction. Journal of Applied Social Psychology. (1999); 29(11): 2308-2327.

Mazzocco MMM, Sonna NL, Shapiro BK, Pinit A, & Reiss AL. Effective procedures for conducting genetic screening studies with children. Journal of Applied Social Psychology. (1998); 28: 23-40.

Mazzocco MMM. Children's interpretations of homonyms: A developmental study. Journal of Child Language. (1997); 24(2): 441-467.

# Mazzocco MMM, Denckla MB, Singer H, Scanlon DM, Vellutino FR, & Reiss AL. Neurogenetic and neurodevelopmental pathways to learning disabilities. Learning Disabilities. (1997); 8(1): 31-42.

# Mazzocco MMM, & Reiss AL. Genetic advances and disorders. Current Opinion in Psychiatry. (1994); 7(5): 392-396.

Mazzocco MMM. The magic of reading aloud. Journal of Youth Services in Libraries. (1993); 6(3): 312-314.

Manuscripts on fragile X syndrome:

Murphy, M. M., & Mazzocco, M. M. M. (2009). The trajectory of mathematics skills and working memory thresholds in girls with fragile X syndrome. Cognitive Development, 24, 430-449.

Dennis, M., Berch, D. B., & Mazzocco, M. M. M. (2009). Mathematical learning disabilities in special populations: Phenotypic variation and cross-disorder comparisons. Developmental Disabilities Research Reviews, 15, 80-89.

Roberts J, Mazzocco MMM, Murphy MM, & Hoehn-Saric R. Arousal modulation in females with fragile X or Turner Syndrome. Journal of Autism and Developmental Disorders (2008), 38: 20-27.

Murphy MM, & Mazzocco MMM. Rote numeric skills may mask underlying mathematical disabilities in girls with fragile X syndrome. Developmental Neurospychology (2008), 33: 345-364.

Mazzocco MMM, & Holden JJA. Fragile X Syndrome. In: I. Brown & M. Percy (Eds.), Intellectual and Developmental Disabilities. (2007); Baltimore, MD: Brookes Publishing. pp 173-187.

Mazzocco MMM, Murphy MM, McCloskey M. The Contribution of Syndrome Research to Understanding Mathematics Learning Disability: The Case of Fragile X and Turner syndromes. In D Berch & MMM Mazzocco (Eds.), Why is Math So Hard for Some Children: The Nature and Origins of Mathematics Learning Difficulties and Disabilities. Brookes Publishers (2007). pp173-193.

Lasker A, Mazzocco MMM, Zee D. Ocular Motor indicators of executive dysfunction in fragile X and Turner syndromes. Brain and Cognition (2007); 63:203-220.

Mazzocco MMM, Thompson L, Sudhalter V, Belser RC, Lesniak-Karpiak K, & Ross JL. Language use in females with fragile X or Turner syndrome during initial social interactions. Journal of Developmental and Behavioral Pediatrics (2006); 27:319-328.

Mazzocco MMM, Bhatia NS, & Lesniak-Karpiak K. Visuospatial skills and their association with math performance in girls with fragile X or Turner Syndrome. Child Neuropsychology (2006); 12:87-100.

Murphy MM, Mazzocco MMM, Gerner G, & Henry AE. Mathematics learning disability in girls with Turner Syndrome or fragile X Syndrome. Brain and Cognition (2006); 6:195-210.

Holden JJA, & Mazzocco MMM. Fragile X Syndrome. To appear in: Brown, (ed). Intellectual and Developmental Disabilities. (in press); Baltimore, MD: Brookes Publishing.

Mazzocco MMM, & McCloskey M. Math performance in girls with Turner or Fragile X syndrome. In Campbell, J (ed), Handbook of Mathematical Cognition. (2005); Hove, East Sussex, Great Britain: Psychology Press. pp 269 -297.

Kirk J, Mazzocco MMM, & Kover, ST. Assessing executive dysfunction in girls with fragile X or Turner syndrome using the Contingency Naming Test. Developmental Neuropsychology. (2005); 28(3): 755-777.

Lesniak-Karpiak K, Mazzocco MMM, Ross JL. Behavioral assessment of social anxiety in females with Turner or fragile X syndrome. Journal of Autism and Developmental Disorders. (2003); 33(1): 55-67.

Keysor CS, Mazzocco MMM, McLeod D, & Hoehn-Saric R. Physiological arousal in females with fragile X or Turner syndrome. Developmental Psychobiology. (2002); 4(2): 133-146.

# Keysor CS, & Mazzocco MMM. A developmental approach to understanding fragile X syndrome in females. Microscopy Research and Techniques. (2002); 57(3): 179-186.

Mazzocco MMM. Math learning disability and math LD subtypes: Evidence from studies of Turner syndrome, fragile X syndrome, and neurofibromatosis type 1. Journal of Learning Disabilities. (2001); 34(6): 520-533.

Myers GF, Mazzocco MMM, Maddalena A, & Reiss AL. No widespread psychological effect of the fragile X premutation in childhood: Evidence from a preliminary controlled study. Journal of Developmental and Behavioral Pediatrics. (2001); 22(6): 353-359.

Mazzocco MMM. Advances in research on the fragile X syndrome. Mental Retardation and Developmental Disabilities Research Reviews. (2000); 6: 96-106.

# Mazzocco MMM. Advances in research on the fragile X syndrome. Mental Retardation and Developmental Disabilities Research Reviews. (2000); 6(2): 96-106.

Teisl JT, Reiss AL, & Mazzocco MMM. Maximizing the sensitivity of a questionnaire for determining at-risk status for fragile X syndrome. American Journal of Medical Genetics. (1999); 83: 281-285.

Mazzocco MMM, & Reiss AL. Size of the FMR2 gene is not associated with normal variation in intelligence. Intelligence. (1999); 27(2): 175-182.

^ Mazzocco MMM, & Reiss AL. A behavioral neurogenetics approach to understanding the fragile X syndrome. In Tager-Flusberg, H (ed.), Neurodevelopmental Disorders: Contributions to a New Framework from the Cognitive Neurosciences. (1999); Cambridge, MA: MIT Press.

Mazzocco MMM, Pulsifer M, Fiumara A, Cocuzza M, Nigro F, Incorpora G, & Barone R. Autistic behaviors among children with fragile X or Rett syndrome. Journal of Autism and Developmental Disorders. (1998); 28(4): 321-338.

Mazzocco MMM, Baumgardner TL, Freund LS, & Reiss AL. Social functioning among girls with fragile X or Turner syndrome and their sisters. Journal of Autism and Developmental Disorders. (1998); 28(6): 509-517.

Mostofsky S, Mazzocco MMM, Aakalu G, Warsofsky IS, Denckla MB, & Reiss AL. Decreased cerebellar posterior vermis size in fragile X syndrome: Association with neurocognitive performance. Neurology. (1998); 50: 121-130.

Mazzocco MMM, Myers G, Hamner JL, Panoscha R, Shapiro BK, & Reiss AL. The prevalence of the FMR1 and FMR2 mutations among preschoolers with language delay. Journal of Pediatrics. (1998); 132: 795-801.

Mazzocco MMM, Kates WR, Baumgardner TL, Freund LS, & Reiss AL. Autistic behaviors among girls with fragile X syndrome. Journal of Autism and Developmental Disorders. (1997); 27(4): 415-435.

Abrams MT, Doheny KF, Mazzocco MMM, Knight SJL, Baumgardner TL, Freund LS, Davies KE, & Reiss AL. Cognitive, behavioral, and neuroanatomical assessment of two unrelated male children expressing FRAXE. American Journal of Medical Genetics (Neuropsychiatric Genetics). (1997); 74: 73-81.

Mazzocco MMM, Sonna NL, Teisl JT, Shah N, Pinit A, Shapiro BK, & Reiss AL. The FMR1 and FMR2 mutations are not common etiologies of academic difficulty in school age children. Journal of Developmental and Behavioral Pediatrics. (1997); 18(6): 22-28.

Mazzocco MMM, & Reiss AL. Normal variation in size of the FMR-1 gene is not associated with variation in intellectual performance. Intelligence. (1997); 24(2): 355-366.

Mazzocco MMM & Holden JA. Neuropsychological profiles of three sisters homozygous for the fragile X premutation. American Journal of Medical Genetics. (1996); 64(2): 323-328.

# Mazzocco MMM, Lachiewicz AM, Kovar CG, Freund LS, Baumgardner TL, & Dykens E: Psychological and emotional studies of individuals with the fragile X full mutation: A workshop summary. Developmental Brain Dysfunction. (1995); 8(4-6): 396-404.

Mazzocco MMM, Freund LS, Baumgardner TL, Forman L, & Reiss AL. Neuropsychological and psychosocial effects of the FMR-1 full mutation: A case report of monozygotic twins discordant for fragile X syndrome. Neuropsychology. (1995); 9: 470-480.

Hagerman RJ, Hull CE, Safanda JF, Carpenter I, Staley LW, O'Connor RA, Seydel C, Mazzocco MMM, Snow K, Thibodeau SN, Kuhl D, Nelson DL, Caskey CT, & Taylor AK. High functioning fragile X males: Demonstration of an unmethylated fully expanded FMR-1 mutation associated with protein expression. American Journal of Medical Genetics. (1994); 4: 298-308.

Dorn MB, Mazzocco MMM, & Hagerman. Behavioral and psychiatric disorders in adult male carriers of fragile X. Journal of the American Academy of Child and Adolescent Psychiatry. (1994); 33(2): 256-264.

Mazzocco MMM, Pennington BP, & Hagerman RJ. Social cognition deficits in females with fragile X. Journal of Autism and Developmental Disorders. (1994); 24(4): 473-485.

Mazzocco MMM, Pennington BP, & Hagerman RJ. The neurocognitive phenotype of female carriers of fragile X: Additional evidence for specificity. Developmental and Behavioral Pediatrics. (1993); 14(5): 328-335.

Mazzocco MMM, & O'Connor R. Fragile X syndrome: A guide for teachers of young children. Young Children. (1993); 49(1): 73-77.

Wilson PG, & Mazzocco MMM. Awareness and knowledge of fragile X syndrome among special educators. Mental Retardation.(1993); 31(4): 221-227.

Staley LW, Hull CE, Mazzocco MMM, Thibodeau SN, Snow K, Wilson VL, Taylor A, McGavran L, Weiner D, Riddle J, O'Connor R, & Hagerman RJ. Molecular clinical correlations in children and adults with fragile X syndrome. American Journal of Diseases of Children. (1993); 147: 723-726.

Mazzocco MMM, Hagerman RJ, & Pennington BF. Problem solving limitations among cytogenetically expressing fragile X women. American Journal of Medical Genetics. (1992); 43: 78-86.

Mazzocco MMM, Pennington BF, Cronister AS, & Hagerman RJ. Specific frontal lobe deficits among women with the fragile X gene. Journal of the American Academy of Child and Adolescent Psychiatry. (1992); 31(6): 1141-1148.

Manuscripts on Turner syndrome:

Mazzocco, M. M. M., & Hanich, L. B. (2010). Math achievement, numerical processing, and executive functions in girls with Turner syndrome: Do girls with Turner syndrome have math learning disability? Learning and Individual Differences. (in press).

Mazzocco, M. M. M. (2009). Mathematical learning disability in girls with Turner syndrome: A challenge to defining MLD and its subtypes. Developmental Disabilities Research Reviews, 15, 35-44.

Dennis, M., Berch, D. B., & Mazzocco, M. M. M. (2009). Mathematical learning disabilities in special populations: Phenotypic variation and cross-disorder comparisons. Developmental Disabilities Research Reviews, 15, 80-89.

Roberts J, Mazzocco MMM, Murphy MM, & Hoehn-Saric R. Arousal modulation in females with fragile X or Turner Syndrome. Journal of Autism and Developmental Disorders (2008), 38: 20-27.

Mazzocco MMM, Murphy MM, McCloskey M. The Contribution of Syndrome Research to Understanding Mathematics Learning Disability: The Case of Fragile X and Turner syndromes. In D Berch & MMM Mazzocco (Eds.), Why is Math So Hard for Some Children: The Nature and Origins of Mathematics Learning Difficulties and Disabilities. Brookes Publishers (2007). pp173-193.

Bondy C. & The Turner Syndrome Consensus Study Groupa. Guidelines for the Care of Girls and Women with Turner Syndrome. Journal of Clinical Endocrinology & Metabolism (2007); 92:10-25.

Lasker A, Mazzocco MMM, Zee D. Ocular Motor indicators of executive dysfunction in fragile X and Turner syndromes. Brain and Cognition (2007); 63:203-220.

Mazzocco MMM. The cognitive phenotype of Turner syndrome: Specific learning disabilities. International Congress Series (2006) 1298C:83-92.

Mazzocco MMM, Thompson L, Sudhalter V, Belser RC, Lesniak-Karpiak K, & Ross JL. Language use in females with fragile X or Turner syndrome during initial social interactions. Journal of Developmental and Behavioral Pediatrics (2006); 27:319-328.

Mazzocco MMM, Bhatia NS, & Lesniak-Karpiak K. Visuospatial skills and their association with math performance in girls with fragile X or Turner Syndrome. Child Neuropsychology (2006); 12: 87-110.

Murphy MM, Mazzocco MMM, Gerner G, & Henry AE. Mathematics learning disability in girls with Turner Syndrome or fragile X Syndrome. Brain and Cognition (2006); 61: 195-210.

Russell H F, Wallis D, Mazzocco MMM, Moshang T, Zackai E, Zinn A, Ross JE, & Muenke M. Increased prevalence of Attention-Deficit/Hyperactivity Disorder (ADHD) in girls with Turner syndrome with no evidence of an imprinting effect for cognitive performance or ADHD. Pediatric Psychology (2006); 31: 945-955.

Kirk J, Mazzocco MMM, & Kover, ST. Assessing executive dysfunction in girls with fragile X or Turner syndrome using the Contingency Naming Test. Developmental Neuropsychology. (2005); 28(3): 755-777.

^ Mazzocco MMM, & McCloskey M. Math performance in girls with Turner or Fragile X syndrome. In Campbell, J (ed), Handbook of Mathematical Cognition. (2005); Hove, East Sussex, Great Britain: Psychology Press. pp 269 -297.

Lesniak-Karpiak K, Mazzocco MMM, Ross JL. Behavioral assessment of social anxiety in females with Turner or fragile X syndrome. Journal of Autism and Developmental Disorders. (2003); 33(1): 55-67.

Keysor CS, Mazzocco MMM, McLeod D, & Hoehn-Saric R. Physiological arousal in females with fragile X or Turner syndrome. Developmental Psychobiology. (2002); 4(2): 133-146.

Mazzocco MMM. Math learning disability and math LD subtypes: Evidence from studies of Turner syndrome, fragile X syndrome, and neurofibromatosis type 1. Journal of Learning Disabilities. (2001); 34(6): 520-533.

Mazzocco MMM. A process approach to describing math difficulties in girls with Turner syndrome. Pediatrics (suppl). (1998); 108: 492-496.

Mazzocco MMM, Baumgardner TL, Freund LS, & Reiss AL. Social functioning among girls with fragile X or Turner syndrome and their sisters. Journal of Autism and Developmental Disorders. (1998); 28(6): 509-517.

Reiss AL, Mazzocco MMM, Greenlaw R, Freund LS, & Ross JL. Neurodevelopmental effects of X monosomy. A Volumetric Imaging Study. Annals of Neurology. (1995); 38: 731-738.

Manuscripts on Neurofibromatosis Type 1:

Mazzocco MMM. Math learning disability and math LD subtypes: Evidence from studies of Turner syndrome, fragile X syndrome, and neurofibromatosis type 1. Journal of Learning Disabilities. (2001); 34(6): 520-533.

Denckla MB, Hofman K, Mazzocco MMM, Melham E, Reiss AL, Bryan RN, Harris EL, Lee J, Cox CS, & Schuerholz LJ. Relationship between T2-weighted hyperintensities (UBOs) and lower IQs in children with Neurofibromatosis-1. American Journal of Medical Genetics (Neuropsychiatric Genetics). (1996); 67(1): 98-102.

Mazzocco MMM, Turner JE, Denckla MB, Hofman KJ, Scanlon DC, & Vellutino FR. Language and reading deficits associated with Neurofibromatosis Type 1: Evidence for a not-so-nonverbal learning disability. Developmental Neuropsychology. (1995); 11(4): 503-522.

Manuscripts on Barth syndrome:

Mazzocco MMM, Henry AE, Kelley RI. Barth Syndrome is associated with a cognitive phenotype. Journal of Developmental and Behavioral Pediatrics (2007); 28:22-30.

Mazzocco M.M.M., & Kelley R. (2001). Preliminary evidence for a cognitive phenotype in Barth syndrome. American Journal of Medical Genetics, 102, 372-378.

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